Abstract
Windthrow, i.e. the felling of trees by wind, occurs continually in forest ecosystems. The uprooting of old trees creates multiple microsites (e.g. pit and mound landscape) that are the main source of soil heterogeneity. To determine the impact of pit and mound landscapes on soil features, a beech forest of the Langa district (Mazandaran province, Northern Iran) was studied. An area of 25 ha was considered for this study wherein three microsites were distinguished, including pit bottom (pit), mount top (mound) and level areas (closed canopy). In this area, 22 uprooted trees were also found. Soil samples were collected at different depths (i.e. 0-15, 15-30 and 30-45 cm) from all microsites and analysed. Our study shows that windthrow events should be considered as an important factor in influencing forest ecosystem, as they affect physical (i.e. density, texture and water content), chemical (i.e. pH, organic C, total N, cation exchange capacity and available nutrients) and biological (i.e. soil microbial respiration and earthworm density/biomass) characteristics of soil, thus resulting in pit and mound microsites that may strongly differ with respect to the closed canopy.